There is no difference. The Mac program just connects to the server of whatever mail provider you use, and automatically gets your mail so you don't have to go to the certain website to get it. (for example, I don't have to get on webmail.vt.edu and sign in, etc...because the Mail program fetches my mail and all I do is open up the mail program). You can also have it so that it is checking on multiple mail accounts, like a gmail account or yahoo account, etc.
Hope that helped.
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